Tree cover in savanna ecosystems is usually regarded as unstable, varying with rainfall, fire, and herbivory [[1], [2], [3] and [4]]. In sub-Saharan Africa, elephants (Loxodonta africana) suppress tree cover, thereby maintaining landscape heterogeneity by promoting tree-grass coexistence. In the absence of elephants, tree encroachment may convert savannas into closed-canopy woodlands [[5] and [6]]; when [...]
